A peatland is defined as any ecosystem with a waterlogged substrate and at least 30 cm of peat (Kivinen and Pakarinen 1981). Once peat has accumulated to this depth the availability of essential nutrients falls sharply. Plant species are no longer rooted in mineral soil, and they must absorb all their mineral nutrients from within a waterlogged, anoxic, peat mass. Nutrients, however, are continualLy locked up within the peat, because ...